国家林业公益性行业科研专项201104009;科技部公益性研究项目2007-04-15;国家野外科学观测研究站项目20080615.Supported by the National Forestry Public Welfare Industry Research Project201104009;the Ministry of Science and Technology Public Welfare Research Project2007-04-15;the National Field Science Observation and Research Station Project20080615