山东省自然科学基金ZR2015CM014;农业产业技术体系CARS-30-ZT-06;山东省”双一流”奖补资金SY-L2017YSTD10.This work was supported by Natural Science Foundation of Shandong ProvinceGrant ZR2015CM014;China”s Agricultural Research Sys-temGrant CARS-30-ZT-06;Funds of Shandong ”Double Tops” ProgramGrant SYL2017YSTD10