现代农业产业技术体系建设专项CARS-29-zp-2;山东省“双一流”建设奖补资金SYL2017YSTD10;国家自然科学基金31501738;长江学者和创新团队发展计划IRT15R42.This work was supported by China Agriculture Research SystemGrant CARS-29-zp-2;Funds of Shandong ”Double Tops” ProgramGrant SYL2017YSTD10;National Natural Science Foundation of ChinaGrant 31501738;the Changjiang Scholars and Innovative Research Team in University ProjectGrant IRT15R42