山东省自然科学基金ZR2013CQ028;青岛市应用基础研究计划16-5-1-54-jch;青岛农业大学博士基金663/1111316.This work was supported by Shandong Province Natural Science FoundationGrant ZR2013CQ028;Qingdao Applied Basic Research PlanGrant 16-5-1-54-jch;Qingdao Agricultural University Doctor FundGrant 663/1111316