上海市种业发展项目”沪农科种字2015第3号、沪农科种字2016第1-1号”和国家大麦青稞产业体系CARS-05。This work was supported by the Project from Shanghai Seed Industry Development ”Shanghai Agricultural Science and Technology No.20153;No.20161-1” and the Project from National Barley Industry Technology SystemGrant CARS-05