国家重点研发计划项目2016YFD0300106;国家现代农业产业技术体系建设专项CARS-02-17;国家公益性行业农业科研专项201503117资助.This study was supported by the National Key Research and Development Program of China2016YFD0300106;the China Agriculture Research SystemCARS-02-17;the Special Fund for Agro-scientific Research in the Public Interest201503117