国家重点研发计划项目2016YFD0300106;北京市农林科学院青年科研基金QNJJ201728;北京市农林科学院院级科技创新团队建设项目JNKYT201603;国家现代农业产业技术体系建设专项CARS-02-11;国家自然科学基金青年科学基金项目31601247资助.This study was supported by the National Key Research and Developing Program of China2016YFD0300106;the Youth Research Fund of the Beijing Academy of Agriculture and Forestry SciencesQNJJ201728;the Innovative Team Construction Project of BAAFSJNKYT201603;the China Agriculture Research SystemCARS-02-11;the National Natural Science Fund Youth Science Fund Project31601247