国家自然科学基金项目31601350;山西省基础研究项目2015021152;国家转基因生物新品种培育重大专项2016ZX08005资助.This study was supported by the National Natural Science Foundation of China31601350;the Shanxi Province Fundamental Research Foundation2015021152;the National Major Project for Developing New GM Crops2016ZX08005