国家科技支撑计划项目2014BAD11B04-2;国家自然科学基金项目30840056, 31171496;山东省现代农业体系花生创新团队首席专家专项SDAIT-04-01资助.This study was supported by the National Key Technology Support Program of China2014BAD11B04-2;the National Nature Science Foundation of China30840056, 31171496;Shandong Modern Agricultural Technology & Industry SystemSDAIT-04-01