国家高技术研究发展计划863计划项目2014AA10A601;农业部引进国际先进农业科学技术计划948计划项目2016-X16;深圳孔雀团队计划20130415095710361;中国农业科学院科技创新工程团队项目资助.This study was supported by the National High Technology Research and Development Program of China 863 Program2014AA10A601;The Program of Introducing International Super Agricultural Science and Technology 948 Program2016-X16;Shenzhen Peacock Plan20130415095710361;the Scientific and Technological Innovation Project of Chinese Academy of Agricultural Sciences