国家自然科学基金项目31171578;黑龙江省高校科技创新团队建设计划项目2011TD005;东北农业大学学科团队建设项目团队1资助. This study was supported by the National Natural Science Foundation of China31171578;Heilongjiang Provincial Science and Technology Innovation Team Building Program for Universities2011TD005;the Northeast Agricultural University Discipline Team Construction ProjectGroup 1