国家公益性行业农业科研专项201203052;浙江省科技厅项目2013C02005, 2013C32023;浙江省自然科学基金项目LZ14C130002, LY15C130002;江苏省现代作物生产协同创新中心项目资助. This study was supported by the China Special Fund for Agro-scientific Research in the Public Interest201203052;the Science and Tech-nology Department of Zhejiang Province2013C02005, 2013C32023;the Natural Science Fundation of Zhejiang ProvinceLZ14C130002, LY15C130002;the Jiangsu Collaborative Innovation Center for Modern Crop Production