国家公益性行业农业科研专项经费项目201203096;国家科技支撑计划项目2014BAD11B03;国家现代农业产业技术体系建设专项NYCYTC-00510;高校自主科技创新基金2013PY001资助.This study was supported by the Special Fund for Agro-Scientific Research in the Public Interest201203096;the National Key Technology R&D Program of China2014BAD11B03;the China Agriculture Research SystemNYCYTC-00510;the Fundamental Research Funds for Central Universities2013PY001