国家自然科学基金项目31370690;国家现代农业产业技术体系建设专项CARS-23;中国农业科学院农业科技创新工程CAAS-ASTIP-2014-TRICAAS资助. This study was supported by the National Natural Science Foundation of China31370690;the China Agriculture Research SystemCARS-23;the Agricultural Science and Technology Innovation Program of CAASCAAS-ASTIP-2014-TRICAAS