国家科技支撑计划项目2012BAD11B01;国家公益性行业农业科研专项201103001, 201503121-05;山东省科技发展计划项目2014GNC111007资助. The study was supported by the National Science and Technology Research Projects of China2012BAD11B01;the Special Fund for Agro-Scientific Research in the Public Interest201103001, 201503121-05;the Shandong Provincial Technology Development Program2014GNC111007