国家自然科学基金项目31260342;国家现代农业产业技术体系建设专项GARS-10;云南省重大科技专项2013ZA007项目资助.This study was supported by the National Natural Science Foundation of China31260342;the China Agriculture Research SystemGARS-10;Specialized Research Fund for Science and Technology Projects of Yunnan Province2013ZA007