国家"十二五"科技支撑计划2012BAD14B18-02;四川省烟草公司重点项目SCYC201504;四川省烟草公司泸州市公司重点项目2013003;四川农业大学"双支计划"基金03571890资助.This study was supported by the National Science and Technology Support Program of 12th Five-Year2012BAD14B18-02;the Key Projects of Sichuan Provincial Tobacco CorporationSCYC201504;the Key Projects of Luzhou Branch of Sichuan Provincial Tobacco Company2013003;the "Double Support Plan" Foundation of Sichuan Agricultural University03571890