湖南省重点研发计划项目2015JC3102;湖南农业大学大学生创新计划项目XCX1504;国家自然科学基金项目31300216资助.
This study was supported by the Key Research Projects of Hunan Province2015JC3102;the College Students Innovation Project of Hunan Agricultural UniversityXCX1504;the National Natural Science Foundation of China31300216