国家自然科学基金项目31460544;江西省农业科学院博士启动基金2012CBS011;国家科技支撑计划项目2012BAD05B05;国家公益性行业科研专项201203030资助。This study was supported by the National Natural Science Foundation of China31460544;the Doctoral Scientific Research Foundation of Jiangxi Academy of Agricultural Sciences2012CBS011;the National Support Program of China2012BAD05B05;the China Special Fund for Agro-scientific Research in the Public Interest201203030