国家自然科学31271635;国家现代农业产业技术体系建设专项CARS-03-1-8;山东省自主创新重大关键技术计划项目2014GJJS0201-1资助。 This study was supported by the National Natural Science Foundation of China31271635;the China Agriculture Research SystemCARS-03-1-8;the Innovation Project for Key Technologies in Shandong Province2014GJJS0201-1