国家“十二五”科技支撑计划项目2015BAD26B01;国家粮食丰产科技工程项目河南省子课题2013BAD07B07;国家公益性行业科研专项201303102资助。This study was supported by the National Science & Technology Pillar Program during the 12th Five-year Plan Period2015BAD26B01;the Henan-topic of the National Grain Bumper Science and Technology Project2013BAD07B07;the Special Fund for Scientific Research in the Public Interest of the Ministry of Agriculture