河北省科技支撑计划项目14226401D;国家科技支撑计划项目2013BAD07B05;国家公益性行业农业科研专项201203100;河北省青年优秀拔尖人才专项基金资助。This study was supported by the Key Technology R&D Program of Hebei Province14226401D;the National Key Technology R&D Pro-gram of China2013BAD07B05;the National Special Fund for Agro-scientific Research in the Public Interest201203100;the Hebei Provincial Fund for Outstanding Young Scholars