国家现代农业产业技术体系建设专项CARS-06-02-02;辽宁省教育厅一般项目L2014271;沈阳农业大学博士后基金项目139173资助。 This study was supported by the China Agriculture Research SystemCARS-06-02-02;the General Fund of Department of Education of Liaoning ProvinceL2014271;the Postdoctoral Fund of Shenyang Agricultural University139173