国家自然科学基金项目31470087;国家科技支撑计划项目2012BAD04B01-07,2012BAD4B02;中国农业科学院科技创新工程项目资助。 This study was supported by the National Natural Science Foundation of China31470087;the National Key Technology R&D Program of China2012BAD04B01-07,2012BAD4B02;Science and Technology Innovation Project of Chinese Academy of Agricultural Sciences