国家重点基础研究发展计划973计划项目2015CB150201;国家自然科学基金项目U1302266,31401412;引进国际先进农业科学技术计划948计划项目2011-G23;国家科技支撑计划项目2013BAD01B03-12;高等学校学科创新引智计划111计划项目B12006资助。This study was supported by the National Basic Research Program of China 973 Program2015CB150201;the National Science Founda-tion of ChinaU1302266 and 31401412;the Program of Introducing International Super Agricultural Science and Technology 948 Program2011-G23;the Key Technologies Research and Development Program of China2013BAD01B03-12;the 111 ProjectB12006