湖南省重点学科建设项目资助2013ZDXK006;中南林业科技大学引进高层次人才科研启动基金项目104-0094Fund of Construction of Key Disciplines of Hunan Province Grant No.2013ZDXK006;Scientific Research Starting Fund for High-level Talent Introduction Program of Central South Forestry University of Science and Technology Grant No.104-0094