中国医学科学院中央级公益性科研院所基本业务费2017PT32020,2018PT32001;中国医学科学院医学与健康科技创新工程项目CAMS-2017-I2M-1-011;国家重点研发计划2016YFC0901500;The Non-profit Central Research Institute Fund of Chinese Academy of Medical Sciences2017PT32020, 2018PT32001;CAMS Innovation Fund for Medical ScienceCAMS-2017-I2M-1-011;National Key Research and Development Program of China2016YFC0901500