国家重点研发计划;中国医学科学院医学与健康科技创新工程;北京市优秀人才培养资助?青年拔尖团队项目;国家自然科学基金;Fund program: National Key Research and Development Program of China 2018YFC1315000, 2016YFC0905400, 2017YFC0907900, 2016YFC1302500;CAMS Innovation Fund for Medical Sciences ;Training Program Foundation for the Talents in Beijing City ;National Natural Science Fund