国家自然科学基金82192901,82192904,82192900,81973125;英国Wellcome Trust212946/Z/18/Z,202922/Z/16/Z,104085/Z/14/Z,088158/Z/09/Z;中国香港Kadoorie Charitable基金82192901, 82192904, 82192900, 81973125;National Natural Science Foundation of China212946/Z/18/Z, 202922/Z/16/Z, 104085/Z/14/Z, 088158/Z/09/Z;Wellcome Trust in United Kingdom;Kadoorie Charitable Foundation in Hong Kong of China