国家自然科学基金81941018;中国香港Kadoorie Charitable基金212946/Z/18/Z,202922/Z/16/Z,104085/Z/14/Z,088158/Z/09/Z;英国Wellcome Trust81941018;National Natural Science Foundation of China212946/Z/18/Z, 202922/Z/16/Z, 104085/Z/14/Z, 088158/Z/09/Z;Kadoorie Charitable Foundation in Hong Kong of China;Wellcome Trust in the UK