国家自然科学基金81373065,81773490;国家重点研发计划“生物安全关键技术研发”重点专项2017YFC1200203;上海市第四轮公共卫生体系建设三年行动计划重点学科项目15GWZK0202National Natural Science Foundation of China81373065,81773490;National Key Research and Development Program of China2017YFC1200203;The Fourth Round of Three-year Public Health Action Plan of Shanghai15GWZK0202