国家自然科学基金面上项目81271779;十二·五传染病重大专项分任务2012ZX10004903?004?002;广东省自然科学基金项目2014A030313725;广州市科技计划项目155700012,201508020248 National Natural Science Fund Project81271779;Major Special Tasks for Infectious Diseases of Twelve. Five Plan2012ZX10004903?004?002;Guangdong Natural Science Foundation Project2014A030313725;Guangzhou Science and Technology Project155700012,201508020248