国家自然科学基金31670538;中央高校基本科研业务费专项资金资助项目JC2015-04,16YBB17;国家林业局野生动植物保护与自然保护区管理司年度项目2015-123Supported by the National Science Foundation of China31670538;Fundamental Research Funds for the Central UniversitiesJC2015-04,16YBB17;Project of Department for Wildlife and Forest Plants Protection,SFA of China2015-123