国家自然科学基金项目31170596,31170374,31370589;福建省种业创新和产业化工程项目2014S1477-4;福建省科技厅重大项目2014N5008资助 Supported by the National Natural Science Foundation of China31170596,31170374,31370589;the Seed Industry Innovation Project of Fujian ProvinceFJZY2014-01;the Major S&T Project of Fujian Province2014N5008