国家自然科学基金项目31170596,31170374,31370589、福建省种业创新和产业化工程项目2014S1477-4和福建省科技厅重大项目2014N5008资助 Supported by the National Natural Science Foundation of China 31170596,31170374,31370589,the Seed Industry Innovation Project of Fujian Province FJZY2014-01,and the Major S&T Project of Fujian Province 2014N5008