国家自然科学基金项目31101196、福建省杰出青年基金项目2015J06006、福建省高等学校新世纪优秀人才支持计划JA14095和现代农业产业技术体系建设专项资金项目CARS-20资助 Supported by the National Natural Science Foundation of China 31101196,the Natural Science Foundation of Fujian Province 2015J06006,the Program for New Century Excellent Talents in Fujian Province University JA14095 and the Special Funds for Modern Agricultural Industry Technology System Construction CARS-20