国家自然科学基金项目41071159、 41371485、国家“十一五”重大科技支撑计划课题2008BADA7B03、湖北省自然科学基金重点项目2014CFA116和湖北省公益性科技研究项目GYS0022资助 Supported by the National Natural Science Foundation of China 41071159,41371485,the Key Sci-tech Project of the Eleventh Five-year Plan of China 2008BADA7B03,the Key Project of Natural Science Foundation of Hubei Province 2014CFA116 and the Hubei Provincial Sci-tech Research Project of Public Welfare GYS0022