云南省应用基础研究计划2011FZ214;现代农业产业技术体系建设专项资金CARS-24;国家自然科学基金项目31101591;the Yunnan Provincial Fund for Applied Basic Researches2011FZ214;the Special Fund for China Modern Agriculture Industry Technology System ConstructionCARS-24;the National Natural Science Foundation Program of PR China31101591