“十三五”国家重点研发计划项目2017YFD0301102,2016YFD0300404;“十二五”国家科技计划项目2013BAD07B07-3;河南省农业科学院自主创新专项基金;the National Key Research and Development Program of the Thirteenth Five-Year Plan2017YFD0301102,2016YFD0300404;the National Key Science and Technology Plan of the twelfth Five-Year Plan2013BAD07B07-3;the Special Fund for Independent Innovation of Henan Academy of Agricultural Sciences