国家自然科学基金项目31500264;山东省农业良种工程重大课题项目LKZ201496-1-3;青岛市民生科技计划农业项目14-2-3-36-nsh;the National Natural Science Foundation of China31500264;the Shandong Provincial Agricultural Elite Varieties ProjectLKZ201496-1-3;the Minsheng Science and Technology Project of Qingdao Agriculture14-2-3-36-nsh