国家自然科学基金项目31501713;国家重点研发计划项目2016YFD0201100;国家现代农业产业技术体系建设资金项目CARS-27;the National Natural Science Foundation of China 31501713 ,the National Key Research and Development
Program of China 2016YFD0201100 ,and the China Modern Agriculture Industry System Construction FoundationCARS-27