国家自然科学基金项目31370497,31500409,31760115;云南省高原湿地创新团队项目2012HC007;云南省科技计划项目2017FD103;西南林业大学科技创新基金项目C16056;the National Natural Science Foundation of China31370497,31500409,31760115;the Yunnan Plateau Wetland Innovation Team 2012HC007 .The Yunnan Science and Technology Project 2017FD103.and the Southwest Forestry University Science and Technology Innovation Fund ProjectC16056