公益性行业农业科研专项201303130;国家自然科学青年基金项目41501314;中央级公益性科研院所基本科研业务费专项IARRP-2014-12资助 This work was supported by the Special Scientific Research Fund of Agricultural Public Welfare Profession of China201303130;the National Natural Science Foundation of China41501314;the Basic Research Fund for the Central Public Welfare Scientific InstitutionIARRP-2014-12