农业部现代小麦产业技术体系项目CARS-3-1-19;山东省自然科学基金项目ZR2016CM34;国家自然科学基金项目31101115,31771717资助 This work was supported by the Technology System in Modem Wheat Industry,Ministry of Agriculture,ChinaCARS-3-1-19;the Natural Science Foundation of Shandong ProvinceZR2016CM34;the National Natural Science Foundation of China31101115,31771717