辽宁省自然科学基金项目20170540483;香港海洋公园保育基金项目AW03.1718;中国海油海洋环境与生态保护公益基金会项目资助 This work was supported by the Liaoning Natural Science Foundation20170540483;the Ocean Park Conservation Foundation,Hong KongAW03.1718;the China National Offshore Oil Corporation Marine Environmental and Ecological Protection Public Welfare Foundation