国家重点研发计划项目2017YFD0201900;农业部长江下游棉花与油菜重点实验室开放课题2017-CR01资助 This work was supported by the National Key Research and Development Plan of China2017YFD0201900;the Open Fund from Key Laboratory of Cotton and Rapeseed in the Lower Reaches of the Yangtze River,Ministry of Agriculture2017-CR01