国家小麦产业技术体系项目CARS-3-1-03;国家公益性行业农业科研专项201203033-2;河南科技大学博士科研启动基金项目13480070资助 This work was supported by the China Agriculture Research SystemCARS-3-1-03;the China Special Fund for Agro-scientific Research in the Public Interest201203033-2;the Henan University of Science and Technology Doctoral Research Launch Fund13480070