国家公益性行业农业科研专项201303125;国家自然科学基金项目51609137;国家留学基金资助项目201308210026;辽宁省教育厅项目2009A630,L2014268资助 This work was supported by the China Special Fund for Agro-scientific Research in the Public Interest201303125;the National Natural Science Foundation of China51609137;the Aboard Scholarship of China Scholarship Council201308210026;the Department of Education of Liaoning Province2009A630,L2014268