国家林业公益性行业科研专项201404204;内蒙古自治区自然科学基金项目2016MS0371;林业科技创新平台项目2015-ZYPT-DW-039资助 This work was supported by the National Forestry Public Welfare Industry Scientific Research Project of China201404204;the Natural Science Foundation of Inner Mongolia Autonomous Region2016MS0371;the Project of Forestry Science and Technology Innovation Platform2015-ZYPT-DW-039