国家自然科学基金项目U1404401;河南省高校科技创新团队支持计划项目161RTSTHN012;河南大学资源与环境研究所项目HD-ZHS-201403;河南大学新兴交叉及特色学科培育项目XXJC20140003资助 This work was supported by the National Natural Science Foundation of ChinaU1404401;Henan Province University Scientific and Technological Innovation Team Support Project161RTSTHN012;Henan University Resource and Environment Research Institute ProjectHD-ZHS-201403;Henan University Emerging Intersect and Characteristic Subject Breeding ProjectXXJC20140003